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There are many differences. The most important difference is that the UK runs on a parliamentary system whereas the US uses a presidential system. The US has a written constitution while the UK does not. The US has a clear separation of Church and State while the US does not. The structural/governmental differences could go on forever.

As for the political atmosphere and values, the two countries also differ. Although the UK has political parties with clear and distinct manifestos (equivalent of a US party's platform), the parties tend to be less antagonistic towards each other. I am mainly referring to relations between the Tories (right) and Labour (left) Maybe the US was like this once, but today is seems that parties are extremely angry with each other.

Another huge difference is that in the UK the Prime Minister has far more power than the president. Because he is part of Parliament, and there is extreme party loyalty, he can reasonably expect to pass most of the legislation he proposes.

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