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Let us remember the original lyrics to the song:

Yankee doodle went to town

Riding on a pony

Stuck a feather in his cap

And called it maccaroni.

So, what is the significance of the pony? None, really; it was just a convenient word to rhyme with maccaroni. At the time those lyrics were composed, it was perfectly normal for people to ride ponies.

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Q: Why does Yankee doodle have someone riding on a pony?
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