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No one knows for sure. But it is strongly suspected that they were murdered by either Richard III or one of his retainers.
The reason for this belief is that when Richard III became the King of England both the princes had more legitimate claims to the throne so Richard had to have them killed to secure his position as king. What is known is they both entered the Tower of London in May/Jun 1483 and were never seen again after Jul 1483.
A final interesting fact is in 1674, some workmen remodelling the Tower of London dug up a wooden box containing two small human skeletons. The bones were found at the foot of a staircase. They were found with "pieces of rag and velvet about them", the velvet indicating that the bodies were those of aristocrats.
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